Revisão de Elbella Evans e gêneros afins (Lepidoptera, Hesperiidae, Pyrrhopyginae)

Revision of Elbella Evans and related genera (Lepidoptera, Hesperiidae, Pyrrhopyginae)

Lepidoptera; Hesperiidae; Elbella; related genera; neotropical

Revisão de Elbella Evans e gêneros afins (Lepidoptera, Hesperiidae, Pyrrhopyginae)¹

Revision of Elbella Evans and related genera (Lepidoptera, Hesperiidae, Pyrrhopyginae)

Olaf H. H. Mielke

Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531-970 Curitiba, Paraná, Brasil. Pesquisador do CNPq

ABSTRACT

A systematic study of the genera Microceris Watson, 1893, Elbella Evans, 1951, Parelbella. gen.n. (type species: Hesperia polyzona [Latreille, 1824]), Pseudocroniades. gen.n. (type species: Ericides (sic) machaon Westwood, [1852]) and Protelbella, gen.n. (type species: Pyrrhopyga (sic) alburna Mabille, 1891), a natural group of the Pyrrhopyginae, here called "Elbella Complex", is presented.

The following are spp.n. and sspp.n.: Elbella intersecta paraensis, Brazil: Pará; Elbella intersecta rufitegula, Brazil: Pernambuco, Bahia, Minas Gerais, Espírito Santo and Rio de Janeiro State; Elbella viriditas boliviana, Bolivia: La Paz; Elbella lampra albociliata, Brazil: São Paulo, Paraná and Santa Catarina: Elbella azeta giffordi, Brazil: Distrito Federal and São Paulo; Elbella patrobas mexicana, Mexico: Chiapas and Quintana Roo; Elbella patrobas evansi, Colombia: Cauca, Caldas and Boyaca; Elbella patrobas tingo, Brazil: Rondonia and Peru: Huanuco and Madre de Dios; Elbella patrobas amazonica, Brazil: Amazonas; Parelbella peruana, Peru: Junin; Pseudocroniades machaon seabrai, Brazil: Rio de Janeiro State; Protelbella alburna brasiliensis, Brazil: Amazonas, Rondonia and Mato Grosso, and French Guiana: Montagne aux Cheveaux.

The following genus is a syn.n.: Hegesippe Evans, 1951 of Elbella Evans, 1951. The following species and subspecies are syn.n.: Pyrrhopyga (sic) semidentata Mabille, 1877 and Elbella intersecta ilona Evans, 1951 of Elbella intersecta intersecta (Herrich-Schäffer, 1869): Pyrrhopyge carriae Bell, 1931 of Elbella viriditas viriditas (Skinner, 1920); Pyrrhopyga (sic) menecrates Mabille, 1878 and Pyrrhopyge margimmiscus Hayward, 1935 of Elbella lampra lampra (Hopffer, 1874); Elbella etna moda Evans, 1951 of Elbella etna Evans, 1951; Jemadia umbrata Mabille & Boullet, 1908 of Elbella patroclus patroclus (Plötz, 1879); Pyrrhopyga (sic) jamina Butler, 1870 and Pyrrhopyga (sic) zimra Hewitson, 1871 of Parelbella ahira ahira (Hewitson, 1857); Elbella zesta Evans, 1951 of Croniades pieria (Hewitson, 1857).

The following are comb.n.: Elbella viriditas viriditas (Skinner, 1920); Elbella viriditas chia Evans, 1951; Elbella mariae molinai (Hayward, 1940): Elbella patroclus acala Evans, 1951; Parelbella polyzona (Latreille, [1824]); Parelbella ahira ahira (Hewitson, 1866); Parelbella ahira extrema (Röber, 1925); Parelbella macleannani (Godman & Salvin, 1893); Pseudocroniades machaon machaon (Westwood, [1852]): Protelbella alburna alburna (Mabille. 1891).

The following are stat.n.: Elbella mariae molinai (Hayward, 1940); Elbella lustra Evans, 1951; Elbella blanda Evans, 1951.

The following is a sp.rev.: Elbella patroclus patroclus (Plötz, 1879).

The following is a nom.rev.: Elbella mariae molinai (Hayward, 1940).

The following are stat.rev.: Elbella miodesmiata (Röber, 1925); Parelbella ahira ahira (Hewitson, 1866); Parelbella macleannani (Godman & Salvin, 1893).

Lectotypes are designated for the following species or variety: Myscelus intersecta Herrich-Schäffer, 1869; Pyrrhopyga (sic) fluminis Butler, 1873; Pyrrhopyga (sic) semidentata Mabille, 1877; Pyrrhopyge lamprus Hopffer, 1874; Pyrrhopyga (sic) menecrates Mabille, 1878; Pyrrhopyga (sic) dulcinea Plöz, 1879; Pyrrhopyga (sic) luteizona Mabille, 1877; Hesperia iphinous Latreille, [1824]; Pyrrhopyga (sic) azeta Hewitson, 1866; Pyrrhopyga (sic) patrobas Hewitson, 1857; Jemadia azeta var. melanina Mabille & Boullet, 1908; Pyrrhopyga (sic) ahira Hewitson, 1866; Pyrrhopyga (sic) jamina Butler, 1870; Pyrrhopyga (sic) zimra Hewitson, 1871; Jemadia macleannani Godman & Salvin, 1893; Ericides (sic) machaon Westwood,[1852] Pyrrhopyga (sic) alburna Mabille, 1891.

Neotypes are designated for the following species: Sarbia amoena Röber, 1925; Pyrrhopyga (sic) othello Plötz, 1879; Mimoniades mimetes Mabille, 1909; Jemadia miodesmiata Röber, 1925; Jemadia umbrata Mabille & Boullet, 1908; Jemadia extrema Röber, 1925.

Key words: Lepidoptera, Hesperiidae, Elbella, related genera, neotropical

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AGRADECIMENTOS. Este estudo não teria sido possível sem a ajuda das pessoas e instituições mencionadas no item "Material e Métodos" e os Drs. A. Zinovjev (Zoological Institute, Russian Academy of Sciences, São Petersburgo, Rússia) pelas informações sobre os tipos de Microceris variicolor e Elbella scylla, Gert Hatschbach (Museu Botânico de Curitiba), Armando Cervi (Departamento de Botânica, Setor de Ciências Biológicas, Universidade Federal do Paraná) pelas identificações das plantas e Robert. K. Robbins, Department of Entomology, National Museum of Natural History, Washington, USA pela permissão para o uso do mapa nas figuras das distribuições geográficas; a eles o meu muito obrigado.

Ao Conselho Nacional de Desenvolvimento Cientifico e Tecnológico (CNPq), à Coordenação de Aperfeiçoamento de Pessoal de Ensino Superior (CAPES), ao Deutscher Akademischer Austauschdienst (DAAD), Bonn, Alemanha, à Royal Society (Londres. Inglaterra), à Royal Geographic Society (Londres, Inglaterra) e ao Programa BIOLAT, Smithsonian Institution, Washington D.C., USA, pelas bolsas e auxílios concedidos, o meu muito obrigado.

Agradeço ainda aos doutores Ângelo B. Machado. Arno Lise. Danúncia Urban. Pe. Jesus S. Moure. Jocélia Grazia, Mima M. Casagrande e Waldemiro Gremski, pela leitura criteriosa e pelas sugestões construtivas que ofereceram.

Aos meus mestres: Drs. Romualdo Ferreira D'Almeida, José Oiticica Filho, Alfredo Rei do Rego Barros e Pe. Jesus Santiago Moure. o meu reconhecimento.

Aos colegas do Departamento de Zoologia, Setor de Ciências Biológicas, Universidade Federal do Paraná, pela contribuição no desenvolvimento deste e outros trabalhos e aos Drs Waldemiro Gremski e Imara Milhoretto, diretor e vice-diretora do Setor de Ciências Biológicas, Universidade Federal do Paraná, pelo apoio, a minha gratidão.

Recebido em 30.III.1994; aceito em 09.IX.1994.

Índice

• ¹
  Austin, G.T.; J.P. Brook & O.H.H. Mielke. 1993. Ants, birds & skippers. Trop. Lep., Gainesville, 4, Suppl. 2:1-11.
• Beattie, J.R. 1976. Rhopalocera Directory. Berkeley, J. B. Indexes.
• Bell, E.L. 1931. Studies in the Pyrrhopyginae, with descriptions of several new species (Lepidoptera, Rhopalocera, Hesperiidae). Jour. N.Y. Ent. Soc. 39:417-491.
• ______. 1932. Studies in the genus Phocides with descriptions of new species (Lepidoptera, Hesperiidae). Trans. Amer. Ent. Soc. 58:169-199.
• ______. 1932. Hesperiidae (Lepidoptera, Rhopalocera) of the Roraima and Duida expeditions, with descriptions of new species. Amer. Mus. Novit. 555:1-16.
• ______. 1933. Studies in the Pyrrhopyginae, with descriptions of new species (Lepidoptera, Rhopalocera, Hesperiidae). Jour. N.Y. Ent. Soc. 41:265-295, 481-528.
• ______. 1934. Studies in the Pyrrhopyginae (Lepidoptera, Rhopalocera). Jour. N. Y. Ent. Soc. 42:393-440.
• ______. 1942. New records and species of Hesperiidae from Mexico (Lep. Hesp.). An. Esc. nac. Cienc. biol., Mexico, 2:455-468.
• ______. 1946. A catalogue of the Hesperioidea of Venezuela. Bol. Ent. venezol., Caracas, 5 (3-4):65-203.
• ______. 1946. Some corrections to the "Studies in the Pyrrhopyginae". Jour. N.Y. Ent. Soc. 54:199-201.
• Biezanko, C.M. 1963. VI Hesperiidae da Zona Sueste do Rio Grande do Sul Grande do Sul. Arq. Ent., sér. (A), Pelotas, 24p.
• Biezanko, C.M. & A. Rufinelli. 1962. Lepidópteros americanos de la coleccion de la catedra de entomologia. Estudio sobre la colección Herborn. Rev. Fac. Agron., Montevideo, 50:119-166.
• Biezanko, CM. & O.H.H. Mielke. 1973. Contribuição ao estudo faunístico dos Hesperiidae americanos. IV Espécies do Rio Grande do Sul, Brasil, com notas taxonômicas e descrições de espécies novas (Lepidoptera). Acta biol. paran., Curitiba, 2 (1-4):51-102.
• Bridges, CH.A. 1983. Lepidoptera: Hesperiidae. Notes on species group names. 1, p. 1-129; 2, p. 1-41; 3, p. 1-62; 4, p. 1-30; 5, p. 1-18. Urbana, edição do autor.
• ______. 1988. Catalogue of Hesperiidae (Lepidoptera: Rhopalocera). 1, p. 1-205; 2, p. 1-67; 3, p. 1-78; 4, p. 1-54; 5, p. 1-20; 6, p. 1-7; App. 1, p. I; App. 2, p. 1-8; Annot. 1, p. 1; Annot. 2, p. 1; Annot. 3, p. 1-4; Annot. 4, p. 1-2; Annot. 5, p. 1-6. Urbana, edição do autor.
• ______. 1988. Catalogue of family-group, and genus-group names (Lepidoptera: Rhopalocera). VI p., Part 1, p. 1-8; Part 2, p. 1-3; Part 3, p. 1-4; Part 4, p. 1-141; Part 5, p. 1-33; Part 6, p. 1-68; Part 7, p. 1-18; Part 8, p. 1-61; Part 9, p. 1-20; Part 10, p. 1-8. Urbana, edição do autor.
• Brown Jr., K.S. 1987. Zoogeografia da região do pantanal mato-grossense. An. Prim. Simp. Rec. Nat. e Soc. Econ. Pantanal, EMBRAPA, Brasília, p. 137-178.
• Brown Jr., K.S. & O.H. Mielke. 1967. Lepidoptera of the Central Brazil Plateau. I. Preliminary List of Rhopalocera (Continued): Lycaenidae, Pieridae, Papilionidae, Hesperiidae. Jour. Lep. Soc. 21 (3):145-168.
• ______. 1968. Lepidoptera of the Central Brazil Plateau. III. Partial list for the Belo Horizonte Area, showing the character of the southeastern "Blend Zone". Jour. Lep. Soc. 22:147-157.
• Butler, A.G. 1869-1874. Lepidoptera exotica or descriptions and illustrations of exotic Lepidoptera. London, E. W. Janson, 190 p.
• ______. 1870. The genera of Hesperidae in the collection of the British Museum. Ent. monthly Mag. 7:55-58, 92-99.
• ______. 1870. Descriptions of some new diurnal Lepidoptera, chietly Hesperiidae. Trans. ent. Soc. London. p. 485-520.
• ______. 1873. Descriptions of new species of Lepidoptera. Cist. Ent. 1:151-177.
• ______. 1877. List of Lepidoptera recently collected by Mr. Walter Davis in Peru, with descriptions of a new genus and several new species. Ann. & Mag. nat. Hist. 20 (4):177-129.
• Campos R.F. 1927. Catalogo preliminar de los lepidopteros del Ecuador. Primera parte Ropaloceros. Rev. Col. Nac. Vicente Rocafuerte, Guayaquil, 9 (27-28):3-106.
• Cardoso, A. 1949. Lepidópteros de Alagoas. Rev. Ent., Rio de Janeiro, 20:427-436.
• Cock. M.J.W. 1984. Lepidoptera notes 1-6. Living World, 1983-1984, p. 35-37.
• Comstock. J.A. & L. Vázquez G. 1961. Estudios de los ciclos biológicos en Lepidópteros mexicanos. An. Inst. Biol. Univ., México. 31:349-448, 54 figs.
• Cresson Jr., E.T. 1926. List of new genera and species described by Henry Skinner. Ent. News 37:246-249.
• Devries, PH.J. 1983. Checklist of Butterflies, p. 654-678. In: D.H. Janzen (ed.). Costa Rican Natural History. Chicago e London, The University of Chicago Press.
• Doubleday, E.; J.O. Westwood & W.C. Hewitson. 1846-1852. The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habits and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson. London, Longman, 534 p.
• Draudt, M. 1921. Hesperidae [não a parte geral], pp. 836-1011, 1046-1055, pls. 113B, 160-193. In A. SEITZ. The Macrolepidoptera of the World 5. Stuttgart, A. Kernen.
• Druce, H.H. 1908. Descriptions of some new Hesperiidae from Central and South America. Trans. ent. Soc. London, p. 375-386.
• Druce, H. 1876. List of the butterflies of Peru, with descriptions of new species. With some notes by Edward Bartlett. Proc. zool. Soc. London, p. 205-250.
• Ebert, H. 1969. On the frequency of butterflies in eastern Brazil, with a list of the butterfly fauna of Poços de Caldas, Minas Gerais. Jour. Lep. Soc. 23, Suppl. 3:1-48.
• Evans, W.H. 1941. An interesting case of development of certain South American Hesperiidae (Lep. Rhopalocera). Proc. roy. ent. Soc. London (A) 16:21-23.
• ______. 1951. A catalogue of the American Hesperiidae, indicating the classification and nomenclature adopted in the British Museum (Natural History). Part I, Pyrrhopyginae. London, British Museum (Natural History), VII+92p.
• Freeman, H.A. 1966. New Hesperiidae records from Mexico. Jour. Lep. Soc. 20 (4):226-228.
• Gibson, W.W. & J.L. Carillo S. 1959. Lista de insetos en la coleccion entomologica de la oficina de estudios especiales, S. A. G. Folleto Misc, Secret. Agrie. Ganad., Oficina Estudios Especiales, Mexico, 9:I-XVIII, 1-254.
• Gillham, N.W. & P.R. Ehrlich. 1955. The butterfly type of Henry Skinner and co-authors in the Academy of Natural Sciences of Philadelphia (Lepidoptera: Papilionoidea and Hesperioidea). Trans. Amer. Ent. Soc. 80:91-117.
• Godman, F.D. 1907. Notes on the american species of Hesperiidae described by Plötz. Ann. & Mag. Nat. Hist. (7) 20:132-155.
• Godman, F.D. & O. Salvin. 1879-1901. Biologia Centrali-Americana. Insecta. Lepidoptera-Rhopalocera. 1 (1879-1886), p. 1-487; 2 (1887-1901), p. 1-782; 3
• ______. 1880. A list of diurnal Lepidoptera collected in the Sierra Nevada of Santa Marta, Colombia, and the vicinity. Trans. ent. Soc. London, p. 119-132.
• Hayward, K.J. 1932. Lepidopteros argentinos. Familia Hesperidae. I Subfamilia Pyrrhopyginae. Rev. Soc. ent. arg. 5 (1):19-35, 2 pls.
• ______. 1934. Lepidopteros argentinos. Família Hesperidae. IV Subfamilia Pamphilinae. V Resúmen, Clave, Apéndices e Indice. Rev. Soc. ent. arg. 6:97-233.
• ______. 1935. Revision de espécies argentinas de Pyrrhopyginae (Lep. Hesp.). Rev. Soc. ent. arg. 7:123-129.
• ______. Some Hesperiidae trom Yungas of Bolivia (Lep.). Rev. Ent., Rio de Janeiro, 8:106-111.
• ______. 1940. A new species of Pyrrhopyge (Lep. Hesp.). Rev. chil. Hist. nat. 44:33-41, fig.
• ______. 1941. Hesperiidarum Argentinae Catalogus. Rev. Mus. La Plata (n. ser.), Zoologia 2:227-340, 1 fig.
• ______. 1942. Notes on Hesperiidae in the collection of the Museu Nacional do Brasil. Bol. Mus. Nac., Rio de Janeiro. 14-17:65-75.
• ______. 1945. Hesperiidae (Lep.) capturados em Porto Cabral durante uma excursão à margem paulista do rio Paraná. Pap. Av. Dep. Zool., São Paulo. 5 (21):197-202.
• ______. 1947. Catalogus Hesperiidarum Rei Publicae Colombianae. Acta zool. lill. 4:201-392.
• ______. 1948. Insecta, Lepidoptera (Rhopalocera), familia Hesperiidarum, subfamiliae Pyrrhopyginarum et Pyrginarum. Gen. Spec. Anim. Argent. 1. Tucuman, Instituto Miguel Lillo, 389p.
• ______. 1973. Catalogo de los ropaloceros argentinos. Op. lill. 23:1-318.
• Hemming, F. 1941. The dates of publication of the several portions of Doubleday (E.) "Genera of Diurnal Lepidoptera" and of the continuation thereot by Westwood (J. O.). Jour. Soc. Bibliogr. nat. Hist., London. 1:335-411.
• ______. 1941. The dates of publication of the specific names first published in Doubleday (E.) "Genera of Diurnal Lepidoptera" and in the continuation thereot by Westwood (J. O.). Jour. Soc. Bibliogr. nat. Hist., London. 1:447-464.
• ______. 1967. The generic names of the butterflies and their type-species (Lepidoptera: Rhopalocera). Bull. Brit. Mus. (Nat. Hist.), Ent., Suppl. 9:1-509.
• Heppner. J.B. & G. Lamas. 1982. Acronyms for World Museum Collections of Insects, with Emphasis on Neotropical Lepidoptera. Bull. Ent. Soc. Amer. 28 (3):305-315.
• Herrich-Schäffer, G.A.W. 1864-1871. Prodromus Systematis Lepidopterorum - Versuch einer systematischen Anordnung der Schmetterlinge. Corr.-Blatt zool.-min. Ver. Regenshurg 18 (1864): 89-112, 123-136, 148-152, 173-181; 19 (1865): 63-76, 84-92, 100-108; 21 (1867): 100-106. 124-128, 138-144, 161-172; 22 (1868): 119-138, 172-176; 23 (1969): 56-64. 67-77, 130-141. 163-172. 184-204; 24 (1870): 154-160; 25 (1871): 103-104.
• Hewitson, W.C. 1851-1877. Illustrations of new species of exotic butterflies, selected chiefly from the collection of W. Wilson Saunders and William C. Hewitson. 5 vols. London. V. Voorst, texto e planchas.
• ______. 1866. Descriptions of new Hesperidae. Trans. ent. Soc. London (3) 2:479-501.
• Hoffmann, C.C. 1940-1941. Catalogo sistemático y zoogeogrático de los lepidópteros mexicanos. Primera parte Papilionoidea. Segunda parte Hesperioidea. An. Inst. Biol., Mexico, 11 (1940):639-739, 2 mapas; 12 (1941):237-294.
• Hoffmann. F. 1934. Beiträge zur Lepidopterenfauna von Sta. Catarina (Süd-brasilien). Ent. Rdsch., Stuttgart, 51:13-15, 25-28, 62-64, 71-74, 248-251, 265-268, 272-277.
• Hopffer, C. 1874. Neue Lepidopteren von Peru. Stett. ent. Ztg. 35:329-371.
• Jong, R. de. 1983. Annotated list of the Hesperiidae (Lepidoptera) of Surinam, with descrition of new taxa. Tijd. Ent. 126:233-268.
• Kaye, W.J. 1915. Pyrrhopygid ova and imagines. Proc. ent. Soc. London 5:CXIX.
• Kirby, W.F. 1871. A Synonymic Catalogue of Diurnal Lepidoptera. London, Voorst, p. VI + 1-690.
• ______. 1877. A Synonymic Catalogue of Diurnal Lepidoptera. Supplement. London, Voorst, p. VI+691-884.
• ______. 1879. Catalogue of the Colleetion of Diurnal Lepidoptera formed by the late William Chapman Hewitson, of Oatlands, Walton-on-Thames, and bequeathed by him to the British Museum. London, Voorst, IV + 246p.
• ______. 1880. Catalogue of the Lepidoptera (Rhopalocera, Sphingidae, Castniidae, and Uraniidae) in the Museum of Science and Art, Dublin, with remarks on new or interesting species. Proc. r. Dublin Soc. (2) 2:292-340.
• Köhler, P. 1923. Fauna argentina. Lepidoptera e collectione Alberto Breyer. I Teil. Rhopalocera. Systematischer Katalog und Studien, Berichtigungen u. Neubeschreibungen. Zeitsehr. wiss. Insektenbiol., Berlin, 18, Sonderheft, 34p.
• Köhler, P. & R. Strassberger. 1928. Catálogo de lepidópteros argentinos. Enumeracion sistemática de lepidópteros diurnos y parte de nocturnos (de Sphingidae hasta Noctuidae (Heliothinae). Buenos Aires, Publ. Breyer, 12p.
• Lamas, G. 1969. Lista de Ropaloceros (Lepidoptera) peruanos citados en la obra "Die Gross-Schmetterlinge der Erde" de Adalbert Seitz. Biota, Lima, 7 (58):265-328; (59):329-354.
• ______. 1981. La fauna de mariposas de la reserva de Tambopata, Madre de Dios, Peru (Lepidoptera, Papilionoidea y Hesperioidea). Rev. Soc. Mex. Lep. 6 (2):23-40.
• ______. 1983. Adiciones y recorrecciones a la lista de mariposas de la Reserva Tambopata. Rev. Soc. Mex. Lep. 8 (1):13-24.
• ______. 1983. Mariposas atraidas por hormigas legionarias en la Reserva de Tambopata, Peru. Rev. Soc. Mex. Lep. 8 (2):49-51.
• Lamas, G.; R.K. Robbins & D. Harvey. 1991. A preliminary survey of the butterfly fauna of Pakitza, Parque Nacional de Manu, Peru, with an extimate of its species richness. Publ. Mus. Hist. nat., UNMSM (A), Lima, 40:1-19.
• Lamas, G.; O.H.H. Mielke & R.K. Robbins. 1993. The Ahrenholz technique for attrackting tropical skippers (Hesperiidae). Jour. Lep. Soc. 47 (1):8 0-82.
• Latreille, P.A. & J.B. Godart. 1819-[1824]. Encyclopédie Methodique. Histoire Naturelle. 9. Entomologie. 828p.
• Lesse, H. de & K.S. Brown Jr. 1971. Formules chromosomiques de Lepidoptères Rhopalocères du Brésil. Bull. Soc. ent. France 76:131-137.
• Lewis, H.L. 1973. Mariposas del mundo. Barcelona, Omega, XVI + 312p.
• Lindsey, A.W. 1925. The Cornell University Expedition to South America of 1919-1920. Scientitic Results. No. 2: Hesperioidea. Denison Univ. Bull., Jour. Sc. Lab. 21:71-1 13.
• ______. 1925. The types of Hesperioid genera. Ann. Ent. Soc. Amer. 18:75-106.
• Llorente-Bousquets. J.; A. Luis-Martínez & I. Vargas-Fernández. 1990. Catalogo sistematico de los Hesperioidea de Mexico 1990. Publ. esp. Mus. Zool., Mexico, 1:1-70.
• Mabilde, A. 1896. Guia pratica para os principiantes coleccionadores de insetos contendo a descripção fiel de perto de 1000 borboletas com 280 figuras lythographadas em tamanho, formas e dezenhos conforme o natural. Porto Alegre, Gundlach & Schludt, 238 p.
• Mabille, P. 1877-1878. Descriptions de Lepidoptéres nouveaux du groupe des Hespérides. Pet. Nouv. Ent. 2:161-162 (1877), 165-166 (1877), 197-198 (1878), 201-202 (1878), 205 (1878), 229-230 (1878), 233-234 (1878), 237-238 (1878), 242 (1878), 261 (1878).
• ______. 1878. Catalogue des Hespérides du Musée Royal D'Histoire Naturelle de Bruxelles. Ann. Soc. ent. Belg. 21:12-44.
• ______. 1891. Description d'Hespérides nouvelles. Ann. Soc. ent. Belg. 35, CR: LIX-LXXXVIII. CVI-CXXI, CLXVIII-CLXXXVII.
• ______. 1903-1904. Lepidoptera Rhopalocera. Fam. Hesperidae, p. 1-78 (1903); 79-210, 4 pls. (1904). In: P. WYTSMAN. Genera Insectorum 17. Bruxelles, Verteneuil & Desmet.
• ______. 1909. Note sur l'Hesperia Iphinous Latr. (Lep. Hesperidae). Bull. Soc. ent. France. p.334-335.
• ______. 1912. Lepidopterorum Catalogus. Pars 9. Hesperiidae; Subfamilia Pyrrhopyginae. Berlin, Junk, 18p.
• Mabille, P. & E. Boullet. 1908. Essai de révision de la famille des Hespérides. Ann. Sc. nat. (Zool.) (9) 7:167-207.
• ______. 1912. Essai de révision de la famille des Hespérides. Ann. Sc. nat. (Zool.) (9) 16:1-159.
• Martin. M. de. 1941-1942. La coleccion de lepidópteros del Museo. Bol Mus. Hist. nat. J. Prado, Lima. 5 (1941): 46-61. 197-208. 343-352. 447-457: 6 (1942) 67-70. 223-228, 343-348.
• Mathew, G.F. 1882. Remarks on some central american species of Pyrrhopyge, Hübn. Ent. monthly Mag. 19:18-19.
• Maza E., J. de la & R. de La maza E. 1985. La fauna de mariposas de Boca de Chajul. Chiapas, Mexico (Rhopalocera). Parte 1. Rev. Soc. Mex. Lep. 9:23-44.
• Maza E., J. de la; A. White L. & R.G. de la Maza E. 1991. La fauna de mariposas de México. Parte 2. Hesperioidea (Lepidoptera: Rhopalocera). Rev. Soc. Mex. Lep. 14:3-44.
• Maza R., R. de la. 1987. Mariposas mexicanas. Guia para su colecta y determinación. Mexico, Fundo de Cultura Económica. 302p.
• Maza E., R.G. de la. 1976. Notas sobre lepidopteros de Rancho Viejo y Tepoztlan, Morelos, Mexico. Segunda parte: Hesperidos, Megatimidos y Castnidos. Rev. Soc. Mex. Lep. 2:15-23.
• Maza E., R.G. de la & A. White L. 1990. Rhopalocera de la Huasteca Potosina, su distribución, composición, origen y evolución. Rev. Soc. Mex. Lep. 13 (2):31-88.
• Maza E., R.G. de la Maza & D.G. Carbonell. 1992. Ropaloceros de Quintana Roo, su distribución, origen y evolucion. Rev. Soc. Mex. Lep. 15 (1):1-43; Anexo 1:1-14.
• Ménétriés, E. 1855-1863. Enumeratio Corporum Animalium Musei Imperialis Academiae Scientiarum Petropolitanae. Classis Insectorum, Ordo Lepidopterorum. Pars 1 (1855), 15+102p.; Pars 2 (1857), 6 + 90p.; Pars 3 (1863), 4 + 20p., Petropoli, Typis Academiae Scientiarum Imperialis.
• Mielke, O.H.H. 1967. Lepidópteros do planalto central brasileiro. IV: Notas complementares sôbre os Hesperiidae da região e descrição de cinco espécies novas (Lepidoptera, Hesperiidae). Rev. Bras. Biol. 27 (2):125.-134.
• ______. 1968. Contribuição ao estudo faunístico dos Hesperiidae brasileiros. I. Resultados de uma excursão à Foz de Iguaçu, Paraná, Brasil, com notas taxonômicas (Lepidoptera). Atas Soc. Biol. Rio de Janeiro 12 (2):73-78.
• ______. 1971. As espécies de Hesperiidae descritas por J. F. Zikán (Lepidoptera). Rev. Bras. Biol. 31 (2):249-262.
• ______. 1980. Contribuição ao estudo faunístico dos Hesperiidae americanos. V. Nota suplementar às espécies de Pyrrhopyginae e Pyrginae do Rio Grande do Sul, Brasil (Lepidoptera). Acta biol. paran. 8-9:7-17.
• ______. 1989. Sobre os tipos de Hesperiidae descritos por Roeber (Lepidoptera). Revta bras. Zool. 6:131-146.
• Mielke, O.H.H. & M.M. Casagrande. 1988. Sobre os tipos de Lepidoptera depositados em museus brasileiros. XX. Pieridae descritos por C.M. Biezanko e H. Fruhstorfer e Hesperiidae por K.T. [recteJ.] Hayward. Revta bras. Ent. 32 (2):115-118.
• Miles Moss, A. 1949. Biological notes on some Hesperiidae of Pará and the Amazon (Lep. Rhop.). Acta zool. IIII. 7:27-79.
• Okano, K. 1981. Studies on the mexican Butterflies (1). Additions and revisions to the Hoffmann's "Catalogo sistematico y zoogeografico de los Lepidópteros mexicanos". Part 1: Hesperiidae (Lepidoptera: Rhopalocera). Tokurana 1:23-139.
• Pallister, J.C 1956. Skippers taken on the Frank C. Johnson Entomological Expedition to Peru, with distribuition and ecological notes. Amer. Mus. Novit. 1763:1-69.
• Plötz, C. 1879. Die Hesperiinen-Gattung Pyrrhopyga (sic) und ihre Arten. Stett. ent. Ztg. 40:520-538.
• Raymond, TH. 1982. Mariposas de Venezuela. Caracas, Corpoven. 275p.
• Riley, N.D. 1926. On the identity of certain Hesperiidae (Lep.) described by Latreille. Trans. ent. Soc. London 74:231-241.
• Röber, J. 1925. Neue siidamerikanische Falter (Lep.). Ent. Mitt., Berlin, 14:85-100, 156-162.
• Shepard, H.H. 1931. Lepidopterorum Catalogus 47. Hesperidae: Subfamilia Pyrginae I. Berlin. Junk. 144p.
• ______. 1936. Lepidopterorum Catalogus 74. Hesperidae: Subfamilia Pyrginae IV. 's-Gravenhage, Junk, p.561-679.
• Silva, A.G. d'A.; C.R. Gonçalves; D.M. Galvão; A.J.L. Gonçalves: J. Gomes; M. do N. Silva & L. de Simoni. 1967-1968. Quarto catálogo dos insetos que vivem nas plantas do Brasil, seus parasitos e predadores. Parte 1 (1-2) (1967) Bibliografia entomológica brasileira, XIII+906p.,; Parte 2 (1) (1968) Insetos, hospedeiros e inimigos naturais, XXVI + 622p.; Parte 2 (2) (1968) Índice de insetos e índice de plantas, VIII +265p., Rio de Janeiro. Laboratório Central de Patologia Vegetal.
• Skinner, H. 1920. Descriptions of new species of Hesperidae. Ent. News 31:132-134.
• Spitz, R. 1932. Catalogo das coleções dos macrolepidopteros do Museu Paulista, com anotações, até 1929. Rev. Mus. paulista 17 (2):833-893.
• Staudinger, O. 1884-1888. Exotische Tagfalter in systematischer Reihenfolge mit Berücksichtigung neuer Arten 1. Fürth, Löwensohn. 333p.
• Steinhauser. S.R. 1975. An annotated list of the Hesperiidae of El Salvador. Bull. Allyn Mus. 29:1-34.
• ______. 1986. A review of the skippers of the narcosius group of species of the genus Astraptes Huebner (sensu Evans, 1952) and erection of a new genus. Lepidoptera: Hesperiidae. Bull. Allyn Mus. 104:1-43, 79 figs.
• Ureta R., E. 1941. Lepidopteros ropaloceros de Bolivia. Bol. Mus. Nac. Hist. nat., Chile, 19:31-41.
• Watson, E.Y. 1893. A proposed classification of the Hesperiidae, with a revision of the genera. Proc. zool. Soc. London, p.3-132.
• Weeks Jr., A.G. 1901. Illustrations of hitherto unfigured Lepidoptera. 32p.
• ______. 1905. Illustrations of diurnal Lepidoptera, with descriptions 1. Boston, Univ. Press, 12+117p.
• ______. 1911. Illustrations of diurnal Lepidoptera 2. Boston, Univ. Press. XVI + 37p.
• Weidemeyer, J.W. 1864. Catalogue of North-American Butterflies. Proc. ent. Soc. Philad. 2:513-542.
• Williams. R.C & K.J. Hayward. 1944. Hesperiidarum Rei Publicae Aequatoris catalogus. Acta zool. IIII. 2:63-246.
• Zikán, J.F. 1922. Unsere Heliconisa-Arten. Dtsch. Ver. Wiss. Kunst, São Paulo, p. 183-193.
• ______. 1927. Biologie der Heliconisa-Arten (Lep. Saturnid.). Dtsch. ent. Ztschr., p.58-82.
• ______. 1928. Die Macro-Lepidopteren des Itatiaya (Südabhang bei Campo Bello). Ent. Rdsch., Stuttgart, 45:7-8. 10-11. 13-14, 19-20, 22-23, 26, 32, 35-36, 38-39, 46.
• ______. 1938. Novos lepidopteros brasileiros da familia Hesperidae. Rev. Ent., Rio de Janeiro, 9 (3-4):321-336.
• Zikán, J.F. & W. Zikán. 1968. Inseto-fauna do Itatiaia e da Mantiqueira. III. Lepidoptera. Pesq. agropec. bras. 3:45-109.

1

Contribuição número 813 do Departamento de Zoologia, Universidade Federal do Paraná. Auxílios ver agradecimentos.

Datas de Publicação

• Publicação nesta coleção
  22 Jul 2009
• Data do Fascículo
  1994

Histórico

• Recebido
  30 Mar 1994
• Aceito
  09 Set 1994